WRKY转录因子主要结合DNA的W-box序列。然而，一些研究发现，WRKY转录因子也可以与非W-box序列结合，如SURE (sugar responsive)元件和TGAC序列等。但是，这方面的研究还处在起步阶段，并不十分明确。
参考文献：
1. Rushton, P. J., Somssich, I. E., Ringler, P., & Shen, Q. J. (2010). WRKY transcription factors. Trends in plant science, 15(5), 247-258.
2. Eulgem, T., Rushton, P. J., Robatzek, S., & Somssich, I. E. (2000). The WRKY superfamily of plant transcription factors. Trends in plant science, 5(5), 199-206.
3. Bakshi, M., & Oelmüller, R. (2014). WRKY transcription factors: Jack of many trades in plants. Plant signaling & behavior, 9(2), e27700.
4. Y. Miao, and R. Zentgraf, A HECT E3 ubiquitin ligase negatively regulates Arabidopsis leaf senescence through degradation of the transcription factor WRKY53, The Plant Journal 63, 179–188 (2010). |
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